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Abbreviations NPs: Nanoparticles Cu: Copper CuO NPs: Copper oxide nanoparticles TM: Total motility PM: Progressive motility VCL: Curvilinear velocity VSL: Straight-line velocity VAP: Average path velocity STR: Straightness LIN: Linearity ALH: Amplitude of lateral head displacement BCF: Beat-cross frequency PMF: Plasma membrane functionality TAC: Total antioxidant capacity MDA: Malondialdehyde CAT: Catalase GSH: Reduced glutathione SOD: Superoxide dismutase AO: Acridine orange ROS: Reactive oxygen species H&E: Hematoxylin and eosin References Auer, J