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Substrate strands (sequences in Supplementary Dataset 1) contained 5 nucleotides of mismatched sequence, allowing differentiation of the two independent strands in each repaired substrate molecule, either by strand-specific fluorescent probes or by next-generation sequencing (see below)

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Transduced cells were selected with 2 g/mL puromycin initially for 2 days, followed by 4 days of puromycin and doxycycline, at that point a T 0 sample was taken